Eras. Essentially the groove got smaller as time passed. Most of the factory '78' cartridges out there are for late stuff (late 40s-50s) so use a 0.25mil (as opposed to the 0.7mil of later mono and more or less all stereo LPs, or 1.0mil of early microgroove mono LPs). Earlier 78s used thicker grooves than this. I have some from the early 20s, and quite a few from the 1930s. Essentially I use the size that's quietest in the groove.